August 6, 2022

The top of molar cusps is seemingly bunoselenomorph

He’s a preliminary and you will upright postprotocrista labially created. New preprotocrista, relatively straight, matches a well-create parastyle. The new metaconule was a bit quicker that is big to the M3. The brand new premetaconulecrista in addition to fossae are pretty straight forward instead of neocrista. Brand new posterior fossa are tilted, particularly on the M3. Out of straight on the M1, the fresh new postmetaconulecrista gets sigmoidal to the M3, skipping the fresh new metacone. The new labial cusps are in an equivalent axis having apparently straight cristae. The paracone labial rib is actually salient, however, as opposed to prior groove. There isn’t any metacone labial rib. This new con el fin de-, meso-, and you will metastyles try salient. The fresh new lingual cingulum is very poor so you can missing such as the entostyle.

The newest registered design of one’s last all the way down premolar is normal getting derived stem pecoran and you can crown pecoran when you look at the that have a highly-create mesolingual conid and you can posterior cristid , . Additionally, the current presence of an enamel socket towards p1 excludes an attribution to help you Prodremotherium otherwise Mosaicomeryx . Even when the pearly whites commonly as bunoselenodont as in Amphitragulus elegans, the new introduced specimens have very ancient possess, such as alt an incredibly quicker metastylid, the lowest top, a tiny postentocristid, and even though laterally compacted, bulged lingual cuspids with the lower molars. Many of these functions ban an affiliation in order to sometimes Dremotherium or Bedenomeryx , . Also, new extremely small-size away from “Amphitragulus” quercyi are an identifying element to the most recent Oligocene ruminants out-of Europe , , , .

Dremotherium cf. guthi from Gaimersheim1: step one remaining maxillary bone on the tooth line M1-M3 (SNSB-BSPG 1952II4855). out of Mailhat: dos left P4-M2 (MNHN LIM154). Babameryx engesseri out of Gaimersheim1: step three remaining fragmentary mandible having m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and you can (c) occlusal viewpoints. The shape pub is actually dos cm.

Holotype known specimens information

Holotype. Jehenne (1987) features designated brand new specimen IPHEP LM.1969.MA.forty as the holotype. It is a limited right maxillary bones that have P3-M3 (, Fig. 2), out-of La Milloque (MP29, France).

Introduced specimens. Mailhat MNHN LIM154 remaining P4-M2 (so it really works, Fig. 6.2); Gaimersheim1 SNSB-BSPG 1952II4855 left maxillary with M1-M3 (that it really works, Fig. six.1); SNSB-BSPG 1952II4870 right m2.


Lower long lasting dentition. The low molar is really poorly kept. Yet not, this huge ruminant and has a totally selenomorph trigonid. This new cristid are elongated therefore the cuspids are large to your enamel quite pleated. The latest metastylid are ripped.

Upper long lasting dentition. The fresh P4 was badly managed. All lingual part try lost. The anterolabial cone possesses a failing labial rib. The fresh new prior stylid is broken. The newest rear one is quick but establish. Zero neocrista blocks the fresh fossa. The fresh new quadratic higher molars into the occlusal glance at happen a properly-establish metaconule quite smaller than the latest protocone. The fresh new cusps are extremely selenomorph. The latest preprotocrista as well as the postmetaconulecrista is actually labially founded and you may some straight. The fresh postprotocrista are much time, very rounded, and labially situated. It could be completely remote (SNSB-BSPG 1952II4855) or bonded (MNHN LIM154) towards premetaconulecrista. Aforementioned ends up within axis of your own mesostyle. This new rear basin can also be features a little metaconule bend for the M2, large with the M3. New paracone, having an excellent paracone rib, in addition to metacone, labially apartment and you can concave, was aligned. New mesostyle and parastyle are very well install. They means round quick columns. The brand new mesostyle is large out-of M1 to M3. The latest entostyle try poor or absent. You will find neither a beneficial lingual cingulum nearby brand new protocone nor good metastyle forming a proper-created cuspid. not, short anterior and you will posterior cingula exist.


So it ruminant is relatively advanced in starting to be better-selenodont and achieving zero lingual cingula when you look at the upper molars, always within Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and you will Babameryx , , , [61–63], . In contrast to Bachitherium, this varieties lacks a keen prior groove with the massive labial paracone rib , , . The newest analyzed fossils be much more much like the cutting-edge stem Pecora and to the latest primitive top Pecora (such as Amphitragulus, Dremotherium, Bedenomeryx). This new tooth is actually quite pleated, whereas it’s fundamentally easy from inside the Prodremotherium. More over, the dimensions of this type of pearly whites is significantly bigger than into the Prodremotherium regarding old Quercy stuff and you can Pech Desse . Tooth be more selenomorph than in “Amphitragulus” and Bedenomeryx , , and the parastyle isn’t that anteriorly projected than noticed with the “Amphitragulus” feningrei , , . Taking a look at the neodiagnosis proposed because of the Mennecart , such fossils are all of the exhibiting features of Dremotherium. The newest currently discussed types into the European countries are D. feignouxi (style of kinds) and D. guthi , , . The latter is actually somewhat smaller compared to the kind variety , similar to the fossils demonstrated right here. D. feignouxi merely taken place for the Agenian (MN1-MN2; ), whereas D. guthi resided into the most recent Oligocene (MP28-MP30; ). Yet not, in lack of more really-managed procedure, a definitive assignation to your species is not possible.